Günther Lab @ Uppsala University

population genomics | ancient DNA | domestication | prehistory


Please check Torsten’s profiles on Google Scholar and ORCID for updated lists and links to all papers.

*=equal contributions


  • Srigyan, M; Bolívar, H; Ureña, I; Sanatana, J; Petersen, A; Iriarte, E; Jakobsson, M; Smith, C; Ibañez, JJ; Götherström, A; Günther*, T; Valdiosera*, C (2020). ‘Bioarchaeological analysis of one of the earliest Islamic burials in the Levant‘, bioRxiv Preprint


  • Schlebusch*, CM; Sjödin*, P; Breton*, G; Günther, T; Naidoo, T; Hollfelder, N; Sjöstrand, AE; Xu, J; Gattepaille, LM; Vicente, M; Scofield, DG; Malmström, H; de Jongh, M; Lombard, M; Soodyall, H; Jakobsson, M (2020). ‘Khoe-San Genomes Reveal Unique Variation and Confirm the Deepest Population Divergence in Homo sapiens’, Molecular Biology and Evolution Journal Link

  • Coutinho, A; Günther, T; Munters, AR; Svensson, EM; Götherström, A; Storå, J; Malmström, H; Jakobsson, M (2020). ‘The Neolithic Pitted Ware culture foragers were culturally but not genetically influenced by the Battle Axe culture herders’, American Journal of Physical Anthropology Journal Link
  • Li, J., Zarzar, T.B.C., White, J.D., Indencleef, K., Hoskens, H., Castrillon, A.O., Nauwelaers, N., Zaidi, A.A., Eller, R.J., Günther, T., Svensson, E.M., Jakobsson, M., Walsh, S., Steen, K.V., Shriver, M.D., Claes, P., (2020). ‘Robust genome-wide ancestry inference for heterogeneous datasets: illustrated using the 1,000 genome project with 3D facial images’, Scientific Reports Journal Link preprint


  • Malmström*, H; Günther*, T; Svensson, EM; Juras, A; Fraser, M; Munters, AR; Pospieszny, Ł; Tõrv, M; Lindström, J; Götherström, A; Storå, J and Jakobsson, M (2019). ‘The genomic ancestry of the Scandinavian Battle Axe Culture people and their relation to the broader Corded Ware horizon’, Proceedings of the Royal Society B  Journal Link
  • Günther, T. & Jakobsson, M. (2019) ‘Population Genomic Analyses of DNA from Ancient Remains’, Book Chapter in Handbook of Statistical Genomics (Fourth Edition), Wiley Publisher Link
  • Günther, T. & Nettelblad C. (2019) ‘The presence and impact of reference bias on population genomic studies of prehistoric human populations’, PLOS Genetics. Journal Link preprint
  • Shebanits K; Günther, T; Johansson, ACV; Maqbool, K; Feuk, L; Jakobsson, M; Larhammar, D (2019). ‘Copy number determination of the gene for the human pancreatic polypeptide receptor NPY4R using read depth analysis and droplet digital PCR’. BMC Biotechnology 19:31. Journal Link
  • Lombard, M; Malmström, H; Schlebusch, C; Svensson, EM; Günther, T; Munters, AR; Coutinho, A; Edlund, H; Zipfel, B; Jakobsson, M (2019) ‘Genetic data and radiocarbon dating question Plovers Lake as a Middle Stone Age hominin-bearing site’, Journal of Human Evolution 131, 203–209. Journal Link
  • Price, N; Hedenstierna-Jonson, C; Zachrisson, T; Kjellström, A; Storå, J; Krzewińska, M; Günther, T; Sobrado, V; Jakobsson, M; Götherström, A; 2019. ‘Viking warrior women? Reassessing Birka chamber grave Bj.581’, Antiquity 93, 181–198. Journal Link


  • Krzewińska, M; Kjellström, A; Günther, T; Hedenstierna-Jonson, C; Zachrisson, T; Omrak, A; Yaka, R; Kılınç, GM; Somel, M; Sobrado, V; Evans, J; Knipper, C; Jakobsson, M; Storå, J; Götherström, A (2018) ‘Genomic and Strontium Isotope Variation Reveal Immigration Patterns in a Viking Age Town’, Current Biology Journal Link
  • Monroy Kuhn, J. M.; Jakobsson, M.; Günther, T. (2018) ‘Estimating genetic kin relationships in prehistoric populations’, PLoS ONE Journal Link preprint
  • Valdiosera*, C; Günther*, T; Vera-Rodríguez, JC; Ureña, I; Iriarte, E; Rodríguez-Varela, R; Simões, LG; Martínez-Sánchez, RM; Svensson, EM; Malmström, H; Rodríguez, L; Bermúdez de Castro, JM; Carbonell, E; Alday, A; Hernández Vera, JA; Götherström, A; Carretero, JM; Arsuaga, JL; Smith, CI; Jakobsson, M (2018) ‘Four millennia of Iberian biomolecular prehistory illustrate the impact of prehistoric migrations at the far end of Eurasia’, Proc Natl Acad Sci U S A Journal Link
  • Günther*, T; Malmström*, H; Svensson*, E; Omrak*, A; Sánchez-Quinto, F; Kılınç, GM; Krzewińska, M; Eriksson, G; Fraser, M; Edlund, H; Munters, AR; Coutinho, A; Simões, LG; Vicente, M; Sjölander, A; Jansen Sellevold, B; Jørgensen, R; Claes, P; Shriver, MD; Valdiosera, C; Netea, MG; Apel, J; Lidén, K; Skar, B; Storå, J; Götherström, A; Jakobsson, M (2018) ‘Population genomics of Mesolithic Scandinavia: Investigating early postglacial migration routes and high-latitude adaptation’, PLoS Biology Journal Link preprint


  • Rodríguez-Varela, R; Günther, T; Krzewińska, M; Storå, J; Gillingwater, TH; MacCallum, M; Arsuaga, JL; Dobney, K; Valdiosera, C; Jakobsson, M; Götherström, A; Girdland-Flink, L (2017) ‘Genomic Analyses of Pre-European Conquest Human Remains from the Canary Islands Reveal Close Affinity to Modern North Africans’, Current Biology Journal Link
  • Schlebusch*, C.M.; Malmström*, H.; Günther, T.; Sjödin, P.; Coutinho, A.; Edlund, H.; Munters, A.R.; Vicente, M.; Steyn, M.; Soodyall, H.; Lombard, M.; Jakobsson, M. (2017) ‘Southern African ancient genomes estimate modern human divergence to 350,000 to 260,000 years ago’, Science Journal Link preprint
  • Hedenstierna-Jonson, C; Kjellström, A; Zachrisson, T; Krzewińska, M; Sobrado, V; Price, N; Günther, T; Jakobsson, M.; Götherström, A; Storå, J; (2017) ‘A female Viking warrior confirmed by genomics’, American Journal of Physical Anthropology
    Journal Link
  • Hollfelder, N; Schlebusch, CM; Günther, T; Babiker, H; Hassan, HY; Jakobsson, M (2017) ‘Northeast African genomic variation shaped by the continuity of indigenous groups and Eurasian migrations’, PLOS Genetics Journal Link
  • Goldberg, A.; Günther, T.; Rosenberg, N.A.; Jakobsson, M. (2017) ‘Ancient X chromosomes reveal contrasting sex bias in Neolithic and Bronze Age Eurasian migrations.’, Proc Natl Acad Sci U S A Journal Link preprint


  • Gattepaille L.; Günther T.; Jakobsson M. (2016) ‘Inferring Past Effective Population Size from Distributions of Coalescent Times.’ Genetics Journal Link
  • Günther, T. & Jakobsson M. (2016) ‘Genes mirror migrations and cultures in prehistoric Europe — a population genomic perspective.’, Current Opinion in Genetics & Development Journal Link preprint
  • Kılınç*, G. M.; Omrak*, A.; Özer*, F.; Günther, T.; Büyükkarakaya, A. M.; Bıçakçı, E.;  Baird, D.; Dönertaş, H. M.; Ghalichi, A.; Yaka, R.; Koptekin, D.; Açan, S. C.; Parvizi, P.; Krzewińska, M.; Daskalaki, E. A.; Yüncü, E.; Dağtaş, N. D.; Fairbairn, A.; Pearson, J.; Mustafaoğlu, G.; Erdal, Y. S.; Çakan, Y. G.; Togan, İ.; Somel, M.; Storå, J.; Jakobsson, M. & Götherström, A. (2016) ‘The Demographic Development of the First Farmers in Anatolia.’, Current Biology Journal Link
  • Günther*, T.; Lampei*, C.; Barilar, I. & Schmid, K. J. (2016) ‘Genomic and phenotypic differentiation of Arabidopsis thaliana along altitudinal gradients in the North Italian Alps.’, Mol Ecol. Journal Link preprint
  • Hervella, M.; Svensson, E. M.; Alberdi, A.; Günther, T.; Izagirre, N.; Munters, A. R.; Alonso, S.; Ioana, M.; Ridiche, F.; Soficaru, A.; Jakobsson, M.; Netea, M. G. & de-la Rua, C. (2016), ‘The mitogenome of a 35,000-year-old Homo sapiens from Europe supports a Palaeolithic back-migration to Africa.’, Sci Rep 6, 25501. Journal Link
  • Omrak, A.; Günther, T.; Valdiosera, C.; Svensson, E. M.; Malmström, H.; Kiesewetter, H.; Aylward, W.; Storå, J.; Jakobsson, M. & Götherström, A. (2016), ‘Genomic Evidence Establishes Anatolia as the Source of the European Neolithic Gene Pool.’, Curr Biol. 26(2), 270–275 Journal Link


  • Günther*, T.; Valdiosera*, C.; Malmström, H.; Urena, I.; Rodriguez-Varela, R.; Sverrisdóttir, Ó. O.; Daskalaki, E. A.; Skoglund, P.; Naidoo, T.; Svensson, E. M.; Bermúdez de Castro, J. M.; Carbonell, E.; Dunn, M.; Storå, J.; Iriarte, E.; Arsuaga, J. L.; Carretero, J.-M.; Götherström, A. & Jakobsson, M. (2015), ‘Ancient genomes link early farmers from Atapuerca in Spain to modern-day Basques.’, Proc Natl Acad Sci U S A 112(38), 11917–11922. Journal Link
  • Gawenda*, I.; Thorwarth*, P.; Günther, T.; Ordon, F. & Schmid, K. J. (2015), ‘Genome-wide association studies in elite varieties of German winter barley using single-marker and haplotype-based methods’, Plant Breeding 134(1), 28–39.


  • Skoglund, P*.; Malmström, H.*; Omrak, A.; Raghavan, M.; Valdiosera, C.; Günther, T.; Hall, P.; Tambets, K.; Parik, J.; Sjögren, K.-G.; Apel, J.; Willerslev, E.; Storå, J.; Götherström, A. & Jakobsson, M. (2014), ‘Genomic diversity and admixture differs for Stone-Age Scandinavian foragers and farmers.’, Science 344(6185), 747–750. Journal Link
  • Siqueira, M. V.; Bonatelli, M. L.; Günther, T.; Gawenda, I.; Schmid, K. J.; Pavinato, V. A. & Veasey, E. A. (2014), ‘Water yam ( Dioscorea alata L. ) diversity pattern in Brazil: an analysis with SSR and morphological markers.’, Genetic Resources and Crop Evolution 61(3), 611–624. Journal Link


  • Günther, T. & Coop, G. (2013), ‘Robust identification of local adaptation from allele frequencies.’, Genetics 195(1), 205–220. Journal Link preprint
  • Günther*, T.; Lampei*, C. & Schmid, K. J. (2013), ‘Mutational Bias and Gene Conversion Affect the Intraspecific Nitrogen Stoichiometry of the Arabidopsis thaliana Transcriptome.’, Molecular Biology and Evolution 30(3), 561–8. Journal Link


  • Hübner, S.; Günther, T.; Flavell, A.; Fridman, E.; Graner, A.; Korol, A. & Schmid, K. J. (2012), ‘Islands and streams: clusters and gene flow in wild barley populations from the Levant.’, Molecular Ecology 21(5), 1115–29. Journal Link


  • Günther, T.; Gawenda, I. & Schmid, K. J. (2011), ‘phenosim – A software to simulate phenotypes for testing in genome-wide association studies.’, BMC Bioinformatics 12(1), 265. Journal Link
  • Günther, T. & Schmid, K. J. (2011), ‘Improved haplotype-based detection of ongoing selective sweeps towards an application in Arabidopsis thaliana.’, BMC Research Notes 4(1), 232. Journal Link
  • Günther*, T.; Schmitt*, A. O.; Bortfeldt, R. H.; Hinney, A.; Hebebrand, J. & Brockmann, G. a. (2011), ‘Where in the genome are significant single nucleotide polymorphisms from genome-wide association studies located?’, OMICS 15(7-8), 507–12. Journal Link
  • Cao*, J.; Schneeberger*, K.; Ossowski*, S.; Günther*, T.; Bender, S.; Fitz, J.; Koenig, D.; Lanz, C.; Stegle, O.; Lippert, C.; Wang, X.; Ott, F.; Müller, J.; Alonso-Blanco, C.; Borgwardt, K.; Schmid, K. J. & Weigel, D. (2011), ‘Whole-genome sequencing of multiple Arabidopsis thaliana populations’, Nature Genetics 43(10), 956–963. Journal Link


  • Günther, T. & Schmid, K. J. (2010), ‘Deleterious amino acid polymorphisms in Arabidopsis thaliana and rice.’, Theoretical and Applied Genetics 121(1), 157–68. Journal Link
  • Childs*, L. H.; Witucka-Wall*, H.; Günther*, T.; Sulpice, R.; V Korff, M.; Stitt, M.; Walther, D.; Schmid, K. J. & Altmann, T. (2010), ‘Single feature polymorphism (SFP)-based selective sweep identification and association mapping of growth-related metabolic traits in Arabidopsis thaliana.’, BMC Genomics 11(1), 188. Journal Link


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